'Auburn' and 'Oregon' are other cultivars (McLeod, 1982). Cv 'Auburn' was selected and developed by the Alabama Agricultural Experiment Station as a forage and cover crop and was certified for use in Alabama and California (Williams et al., 1959).
'Lana' was the most recent successful vetch introduced to California (Miller et al., 1989).
Woollypod vetch produces a moderate proportion of hard seed. The seeds are more oval than are the nearly round seeds of hairy vetch (Williams et al., 1959).
Williams et al. (1959) stated that woollypod vetch is similar in general appearance to hairy vetch; however, the flowers are a little smaller. Woollypod vetch is self-fertile, although very attractive to bees.
This form requires 10 inches or more annual rainfall and is restricted to elevations below 4,000 feet (Williams et al., 1959). The elevational limit was repeated by Graves et al. (1986) and Miller et al. (1989). Finch and Sharp (1983) wrote that the form is well adapted in California below 3,000 ft.
'Lana' is resistant to waterlogging (Miller, 1988).
Stivers & Shennan (pers. comm.) found that cv 'Lana' under normal rainy winter conditions dried out the soil profile down to a depth of 24 inches relative to a winter fallow. The actual amount of water lost in this drying out of the profile was less than 5 percent of the standard amount of irrigation water used to irrigate the summer cash crop.
On rangelands, fertilizing at time of planting with 200 to 500 lbs/acre of single superphosphate will suffice for the majority of range soils. In areas where sulfur is lacking, 200 to 400 lbs. of gypsum per acre is adequate. Refertilize every two or three years (Williams et al., 1959).
Cv 'Lana' was said to tolerate loam to clay soils (Munoz & Graves, 1988), and to be well adapted to all orchard and vineyard soils in California below 3000 ft (Finch and Sharp, 1983).
Woollypod vetch has been grown for seed in California and Western Oregon (McLeod, 1982). Finch and Sharp (1983) noted that cv 'Lana' requires a regrowth period (without mowing) of about 4 weeks before the late May maturity date in order to produce seed for the next year. This was reiterated by Graves et al. (1986).
'Lana' can maintain self-reseeding stands (Munoz & Graves, 1988).
In Syria, rainfed woollypod vetch yielded 1.26 and 1.14 Mg/ha of seeds in separate trials (Abd El Monem et al., 1990).
Abd El Monem et al. (1990) found that in Syria, rainfed woollypod vetch attained maximum biomass yield at 20-50% podding. By contrast, common vetch attained maximum biomass at maturity.
Based on replicated trials in Mendocino wine vineyards, cv 'Lana' shows a great tendency to climb (much greater than for purple or common vetch) and is, therefore, probably unsuitable in most vineyards (Bugg, pers. comm.).
Kutschera (1960) reported that woollypod vetch generally roots to a depth of 80 cm.
Williams et al. (1959) recommended that on rangelands where there is competition from annual grasses or sprouting brush, newly planted woollypod vetch stands should be grazed as soon as competitors are of pasturable height. Grazing should be to a uniform height of about 3 inches and should last no more than 1 month. Avoid grazing while the soil is saturated, or trampling damage may occur. Remove stock well before spring moisture depletion to allow the vetch to regrow and mature a seed crop. A weed-free brush burn planting becomes better established if stock are kept off the first season until seed is ripe.
In a Pennsylvanian trial, 'Lana' germinated well, but spring regrowth was slow and suppressed by weeds (Hofstetter, 1988).
Cv 'Lana' reseeds well (Finch and Sharp, 1983), but under no-till management in an organic vineyard in Mendocino County, California, Bugg et al. (unpublished data) found little 2nd-year regeneration after a heavy seed crop the first year. This may relate to hardseededness of the species.
In his organic walnut orchards, Russell Lester of Solano County California (near Winters, Yolo County) is experimenting (1990-1992) with strip management of 'Lana' vetch and other cover crops. In theory, 10% of the orchard floor devoted to a reseeding remnant strip could more than suffice for reseeding the entire floor to cv 'Lana.' Lester uses flail chopping in late April over most understory vegetation. Remnant strips may have to be shifted from year to year to distribute the seed bank (Bugg, pers. comm.).
Miller et al. (1989) cautioned that cv 'Lana' is not tolerant of frequent mowing and can only be mowed down to a height of five inches. Where frequent, close mowing of the covercrop is needed to reduce the threat of frost, 'Lana' vetch will not reseed.
Cv 'Lana' can be mowed after seed production, if reseeding is desired (Munoz & Graves, 1988).
Cv 'Lana' used as a living mulch should be mowed at the time of planting the vegetable (Munoz & Graves, 1988).
Weedy fields of a high 'Lana' vetch can be mowed or grazed very close to the ground for weed control. The vetch will recover and may be quite competitive with remaining weeds. (Mark van Horn, pers. comm.)
Williams et al. (1959) suggested that, as a cover crop, it may substitute for purple vetch, as it has outperformed the latter in several trials involving planting at a wide row spacing. The relative price of seed may influence the choice.
Woollypod vetch is the preferred vetch for range seeding purposes; 'Lana' vetch offers the best nutritive quality and year-to-year persistence (Murphy et al., 1976). It makes good hay either alone or in mixture with a cereal (Williams et al., 1959).
Vetches are useful as a source of spring feed but even more important as summer dry forage; also, their seeds are fed upon by upland game birds (Murphy et al., 1976).
Cv 'Lana' is suitable in orchards and vineyards (Finch and Sharp, 1983), where a large amount of biomass is needed, but not where more than three mowings are used before April 1 (Finch and Sharp, 1983). It is also very viney and may cause problems in vineyards (Munoz & Graves, 1988).
Williams et al. (1959) wrote that when grown for hay, oat is often suggested as an intercrop for vetch because the former supports the latter and facilitates harvesting. The oat variety 'Kanota' begins flowering at the same time the woollypod vetch is in late bloom, and is therefore a desirable companion. These are the most nutritious stages of growth for hay production. Where hay containing dough-stage oat is desired with woollypod vetch, the variety 'Indio' is suggested. The suggested planting rate is 30 lbs. each per acre each of vetch and oat.
As related by Schenk and Werner (1991), various legumes in the tribe Vicieae (peas, lentils, and vetches) contain Beta-(3-isoxazolinonyl) alanine, which is released into soil as a root exudate and apparently is an allelopathic compound. This chemical can cause reduced growth in seedlings of various grasses and of lettuce. Pea was only slightly affected.
In rangeland plantings, woollypod vetch is compatible for planting in mixture with the annual grasses, 'Blando' brome, and annual ryegrass. They may be added singly or in mixture at the rate of 4 lbs. of grass seed per acre (Williams et al., 1959).
Abd El Monem et al. (1990) found that in Syria, rainfed woollypod vetch attained maximum biomass yield at 20-50% podding. By contrast, common vetch attained maximum biomass at maturity. Rainfed woollypod vetch (Vicia villosa ssp. dasycarpa) at 102, 118 132, and 145 days after planting (D.A.P.) yielded 2.08, 4.01, 4.34, and 3.69 Mg/ha of dry matter during 1986- 87, and during 1987-88 at 115, 130, 144, and 163 D.A.P. the corresponding figures were 2.58, 4.51, 5.88, and 5.68. To summarize, maximum biomass yields were obtained around 132-144 D.A.P, and those yields were 4.34 and 5.68 Mg/ha (Abd El Monem et al., 1990).
At plowdown, of cv 'Lana,' available N (in lbs of N/acre) can be estimated by multiplying harvested fresh weight of cover crop from a sixteen square feet (4 x 4 ft.) sample plot by 18 (Miller et al., 1989).
Hu et al. (1997) conducted a replicated field study at the Student Experimental Farm (SEF) and the Sustainable Agriculture Farming Systems (SAFS) Project, U.C. Davis, CA. The study compared carbon and nitrogen transformations following cover crop incorporation on organically vs. conventionally managed Yolo sandy loam (at SEF: coarse-loamy, mixed, nonacid, thermic Mollic Xerofluvent) and Reiff sandy loam soil (at SAFS: coarse-loamy, mixed, nonacid, thermic Mollic Xerofluvent) Chemical characteristics for the oat and woollypod vetch harvested on April 14 and used as cover crops were as follows (±S.E.M., where indicated):
Cover Crop C/N C N Cellulose LigninPrior to incorporation of cover crops, soil organic N and soil organic C were significantly higher in organically managed than in conventionally managed plots, both at SEF and at SAFS. These differences were lessened or obscured during the 35 days following incorporation. Microbial biomass C was initially greater under organic management at both sites; again, differences were obscured following cover crop incorporation. Cover crop debris buried in litter bags in the SAFS plots showed more rapid disappearance in organic than in conventional plots.Oat 33.6 412±3 g/kg 12.3±0.4 g/kg 349±2.2 g/kg 43±0.8 g/kg Woollypod Vetch 13.3 427±3 g/kg 32±0.9 g/kg 288±3.7 g/kg 84±1.1 g/kg
Hu et al. (1997) conducted a replicated field study at the Student Experimental Farm (SEF) and the Sustainable Agriculture Farming Systems (SAFS) Project, U.C. Davis, CA. The study compared carbon and nitrogen transformations following cover crop incorporation on organically vs. conventionally managed Yolo sandy loam (at SEF: coarse-loamy, mixed, nonacid, thermic Mollic Xerofluvent) and Reiff sandy loam soil (at SAFS: coarse-loamy, mixed, nonacid, thermic Mollic Xerofluvent) Chemical characteristics for the oat and woollypod vetch harvested on April 14 and used as cover crops were as follows (±S.E.M., where indicated):
Cover Crop C/N C N Cellulose LigninPrior to incorporation of cover crops, soil organic N and soil organic C were significantly higher in organically managed than in conventionally managed plots, both at SEF and at SAFS. These differences were lessened or obscured during the 35 days following incorporation. Microbial biomass C was initially greater under organic management at both sites; again, differences were obscured following cover crop incorporation. Cover crop debris buried in litter bags in the SAFS plots showed more rapid disappearance in organic than in conventional plots.Oat 33.6 412±3 g/kg 12.3±0.4 g/kg 349±2.2 g/kg 43±0.8 g/kg Woollypod 13.3 427±3 g/kg 32±0.9 g/kg 288±3.7 g/kg 84±1.1 g/kg Vetch
To ensure high protein feed in summer and early fall, postpone grazing of 'Lana' until after the growing season (Murphy et al., 1976).
Bugg (pers. comm.) noted that pea aphid (Acyrthosiphon pisum) and associated lady beetles (mainly Hippodamia convergens) and lacewings (mainly Chrysoperla carnea) occur on 'Lana' from mid April until maturity; flower thrips (Frankliniella spp.) and associated minute pirate bug (Orius tristicolor) are abundant in flowers. Woollypod vetch lacks extrafloral (stipular) nectaries so is not very attractive to parasitic wasps. Lygus bugs can be abundant.
Bugg et al.(unpublished data) conducted a replicated study (r=4) at Blue Heron Vineyard (Fetzer Vineyards), Hopland, Mendocino County, California. Cover crops were seeded in late October, 1990 and harvested on May 15-16, 1991. Dominant winter annual weeds were chickweed, shepherds purse, rattail fescue, and annual ryegrass. Plots seeded to cv 'Lana' showed no weed biomass production at all; of the 32 cover crops seeded, this was the only entry with that distinction. Vegetational cover by 'Lana' in early May was 97.50+/-2.50 % (Mean +/- S.E.M.).
Various legumes in the tribe Vicieae (peas, lentils, and vetches) contain Beta-(3-isoxazolinonyl) alanine, which is released into soil as a root exudate and apparently is an allelopathic compound. This chemical can cause reduced growth in seedlings of various grasses and of lettuce. Pea was only slightly affected (Schenk and Werner, 1991).

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